Energetic electron and proton fluxes measured by geosynchronous satellites often show sawtooth-like variations during magnetic storms. We examine whether the sawtooth oscillations and relevant magnetospheric-ionospheric disturbances are caused by periodic substorms or driven by a series of enhancements in the solar wind pressure. We show that there are significant differences between periodic substorms and solar wind-induced variations. The energetic fluxes at geosynchronous orbit may increase by orders of magnitude after each onset of periodic substorms and by 10--50% in response to a large solar wind pressure impulse. The sudden increases of the energetic fluxes during periodic substorms show significant time delays of 30--50 min at different longitudes/local times, indicating that the fluxes are injected on the nightside and then drift to the dayside. In contrast, the small flux increases caused by solar wind pressure enhancements occur almost simultaneously at all local times. The periodic substorms always have a strong spectrum peak at 2--3 hours, no matter whether the solar wind pressure and/or IMF have similar spectrum peaks. The nightside magnetospheric magnetic elevation angle shows a large (30--60¿) increase at each onset of periodic substorms through dipolarization and a small (<10¿) decrease in response to a solar wind pressure impulse. Each cycle of periodic substorms can cause a deviation of 40--60 nT in the midlatitude geomagnetic field; the midlatitude geomagnetic deviations caused by solar wind pressure enhancements are proportional to the square root of the pressure change. The increase of the polar cap index caused by substorms is ~4 times that caused by solar wind pressure enhancements. We conclude that the sawtooth-like flux oscillations represent flux injections during periodic substorms and that the period of substorms is determined by the magnetosphere.